The rapid expansion of human populations in East Africa increases human-wildlife interactions, particularly along borders of protected areas (PAs). This development calls for a better understanding of how humanmodified landscapes facilitate or exclude wildlife in savannas and whether these effects change through time. Here, we used camera traps to compare the distribution of 13 large herbivore species in Serengeti National Park with adjacent village lands used by livestock and people at both seasonal and diel cycle scales. The results show