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A literature review of horn and horn-like structures in vertebrates to correlate placement to function, behavior, and niche.

In this paper, the true horns of bovids are compared to the many horn-like structures found throughout the animal kingdom. Literary sources were reviewed for a variety of horn-like structures. The review was limited to extinct and extant terrestrial vertebrates. 3D models of these organisms’ skulls and “horns” were created or acquired and 3D printed at the same scale to illustrate their differences and similarities and to analysis of their comparative size easier between large and small genera.

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Horns and other bony structures of the skull of the giraffe, and their functional significance

The gross morphology of the giraffe skull and its superficial anatomy is described. The parietal, median, occipital, orbital, azygous horns and other exostoses are described. The giraffe’s peculiar mode of intra-specific fighting is explained and the features of the giraffe’s skull are related to this. It is concluded that the skull of the male giraffe is highly specialised for its particular mode of fighting. The male differs in this respect from the female whose skull is not so developed and

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