Maternal investment and maternal care provided by mammalian females can change during their ontogeny. Differences may be related to the age and/or experience (parity) of the female. Several hypotheses have been postulated to explain changes in the rate of maternal care of females during their lifetime. The residual reproductive value hypothesis supposes a higher rate of maternal care in older and more experienced females, in contrast the targeted reproductive effort hypothesis predicts lower levels of maternal care in older and/or