Applying principles of foraging ecology to zoo-housed animals can positively influence animal behavior, and assist with evaluating exhibit space and design. In this study, we implemented zoo foraging ecology by measuring giving-up densities (GUDs) in food patches to address several welfare-related questions with captive okapi (Okapia johnstoni). Our objectives were to: (1) determine whether food patches can reveal how the individual animals perceive their exhibit space (i.e., areas of preference and aversion; landscapes of comfort); and (2) determine whether implementation